Project Statistics: This research was partially supported by Direcção Regional de Florestas, Fundo Florestal Permanente and Autoridade Florestal Nacional, through a national project ‘O nemátode-da-madeira-do-pinheiro (NMP), Bursaphelenchus xylophilus’, by FEDER funds through the Programa Operacional Factores de Competitividade – COMPETE and by national funds through the Fundação para a Ciência e a Tecnologia (FCT), Portugal, under the project PTDC/AGR-CFL/115373/2009. This work was further supported by COST Action FP1305 - Biolink: Linking belowground biodiversity and ecosystem function in European forests. D.N.P. was supported by FCT, postdoctoral fellowship SFRH/BPD/100721/2014.

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