Project Statistics: We thank the Natural Environment Research Council for funding through a studentship grant NE/L501669/1 to S.J.B. C.M.B. acknowledges funding via an ERC Starter Grant ERC-2013-StG-335891-ECOFLAM. S.P.H., T.M.L. and C.M.B. acknowledge funding from the NERC ‘JET’ grant NE/N018508/1, as well as a Royal Society Wolfson Research Merit Award supporting T.M.L.

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