This study was supported by Fundação para a Ciência e a Tecnologia (FCT/MCTES) through national funds UID/AMB/50017/2019 granted to Centro de Estudos do Ambiente e do Mar (CESAM) and UID/ MAR/04292/2019 granted to Marine and Environmental Sciences Centre (MARE), and through the Regional Operational Programme CENTRO2020 within the scope of the project CENTRO-01- 0145FEDER-000006. A.M.M. Gonçalves acknowledges FCT for the financial support provided through the post-doctoral grant (SFRH/BPD/ 97210/2013) co-funded by the Human Potential Operational Programme (National Strategic Reference Framework 2007–2013), European Social Fund (EU), and the program POPH/FSE, and University of Coimbra for the contract (IT057-18-7253). N. Abrantes is recipient of an FCT researcher contract (IF/01198/2014) and M.O. Rodrigues is recipient of a research grant from FCT (SFRH/BD/136931/2018) funded by National Funds and Community Funds through FSE.